species richness, Ecosystems, Ecosystem services, Forests, Habitat, nutrients, Biomass, Laboratories, Livestock, Coleoptera, Grazing, functional traits, Mesocosms, Pasture, coprophagous beetles, ecosystem process, sheep and cattle dung, carabaeinae, Scarabaeidae, Mexican Transition Zone, Manure, Phanaeus, Dichotomius, Dung, Scab